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Journal articles (request a pdf) Simmons, L.W. 1986. Intermale competition and mating success in the field cricket, Gryllus bimaculatus (de Geer). Anim. Behav., 34, 567-579.
Simmons, L.W. 1986. Female choice in the field cricket, Gryllus bimaculatus (de Geer). Anim. Behav., 34, 1463-1470.
Simmons, L.W. 1987. Heritability of of male character chosen by females of the field cricket, Gryllus bimaculatus. Behav. Ecol. Sociobiol., 21, 197-202.
Simmons, L.W. 1987. Sperm competition as a mechanism of female choice in the field cricket, Gryllus bimaculatus. Behav. Ecol. Sociobiol., 21, 197-202.
Simmons, L.W. 1987. Competition between larvae of the field cricket, Gryllus bimaculatus (Orthoptera: Gryllidae) and its effects on some life history components of fitness. J. Anim. Ecol., 56, 1015-1027.
Simmons, L.W. 1987. Female choice contributes to offspring fitness in the field cricket, Gryllus bimaculatus (de Geer). Behav. Ecol. Sociobiol., 21, 313-321.
Simmons, L.W. 1988. The contribution of multiple mating to the lifetime reproductive success of female feild crickets (Gryllus bimaculatus ). Ecol. Entomol., 13, 57-69.
Simmons, L.W. 1988. Male size, mating potential and lifetime reproductive success in the field cricket, Gryllus bimaculatus (de Geer). Anim. Behav., 36, 372-379.
Simmons, L.W. 1988. The calling song of the field cricket, Gryllus bimaculatus (de Geer): constraints on transmission and its role in intermale competition and female choice. Anim. Behav., 36, 380-394.
Simmons, L.W. 1989. Kin-recognition and its influence on mating preferences of the field cricket, Gryllus bimaculatus (de Geer). Anim. Behav., 37, 68-77.
Simmons, L.W. & Parker, G.A. 1989. Nuptial feeding in insects: paternal investment versus mating effort. Ethology, 81, 332-343.
Parker, G.A. & Simmons, L.W. 1989. Nuptial feeding in insects: theoretical models of male and female interests. Ethology, 82, 3-26.
Simmons, L.W. 1990. Post-copulatory guarding, female choice and the levels of gregarine infections in the field cricket, Gryllus bimaculatus. Behav. Ecol. Sociobiol., 26, 403-407.
Simmons, L.W. 1990. Nuptial feeding in tettigoniids: male costs and the rates of fecundity increase. Behav. Ecol. Sociobiol., 27, 43-47.
Simmons, L.W. 1990. Pheromonal cues for the recognition of kin by female field crickets, Gryllus bimaculatus. Anim. Behav., 40, 192-195.
Simmons, L.W. & Bailey, W.J. 1990. Resource influenced sex roles of zaprochiline tettigoniids. Evolution, 44, 1853-1868.
Gwynne, D.T. & Simmons, L.W. 1990. Experimental manipulation of courtship roles in an insect. Nature, 346, 172-174.
Parker, G.A., Simmons, L.W. & Kirk, H. 1990. Analysing sperm competition data: simple models for predicting mechanisms. Behav. Ecol. Sociobiol., 27, 55-65.
Ward, P.I. & Simmons, L.W. 1990. Short-term changes in the numbers of the yellow dung fly, Scathophaga stercoraria (Diptera: Scathophagidae). Ecol. Entomol., 15, 115-118.
Simmons, L.W. 1991. Female choice and the relatedness of mates in the field cricket, Gryllus bimaculatus. Anim. Behav., 41, 493-501.
Simmons, L.W. 1991. On the post-copulatory guarding behaviour of male field crickets. Anim. Behav., 42, 504-505.
Simmons, L.W. & Gwynne, D.T. 1991. The refractory period of female katydids (Orthoptera: Tettigoniidae): sexual conflict over the remating interval ? Behav. Ecol., 2, 276-282.
Simmons, L.W. & Ward, P.I. 1991. The heritability of sexually dimorphic traits in the yellow dung fly, Scathophaga stercoraria (L.). J. Evol. Biol., 4, 593-601.
Parker, G.A. & Simmons, L.W. 1991. A model of constant random sperm displacement during mating: evidence from Scatophaga. Proc. Roy. Soc. Lond. B, 246, 107-115.
Bailey, W.J. & Simmons, L.W. 1991. Male-male behavior and sexual dimorphism of the ear of a zaprochiline tettigoniid (Orthoptera: Tettigoniidae). J. Insect Behav., 4, 51-65.
Ward, P.I. & Simmons, L.W. 1991. Copula duration and testes size in the yellow dung fly Scathophaga stercoraria (L.): the effects of diet, body size and mating history. Behav. Ecol. Sociobiol., 29, 77-85.
Simmons, L.W. 1992. Quantification of role reversal in relative parental investment in a bushcricket. Nature, 358, 61-63.
Simmons, L.W. 1992. Sexual selection and body size in a natural population of the field cricket, Gryllus campestris. J. Orthop. Res., 1, 12-13.
Simmons, L.W. & Parker, G.A. 1992. Individual variation in sperm competition success of yellow dung flies, Scatophaga stercoraria. Evolution, 46, 366-375.
Simmons, L.W. & Zuk, M. 1992. Variability in call structure and pairing success of male field crickets, Gryllus bimaculatus: the effects of age, size and parasite load. Anim. Behav., 44, 1145-1152.
Simmons, L.W., Teale, R.J., Maier, M., Standish, R.J., Bailey, W.J. & Withers, P.C. 1992. Some costs of reproduction for male bushcrickets, Requena verticalis (Orthoptera: Tettigoniidae): allocating resources to mate attraction and nuptial feeding. Behav. Ecol. Sociobiol., 31, 57-62.
Simmons, L.W. 1993. Some constraints on reproduction for male bushcrickets, Requena verticalis (Orthoptera: Tettigoniidae): diet, size and parasite load. Behav. Ecol. Sociobiol., 32, 135-140.
Simmons, L.W. & Bailey, W.J. 1993. Agonistic communication between males of a zaprochiline katydid (Orthoptera: Tettigoniidae). Behav. Ecol., 4, 364-368.
Simmons, L.W. & Gwynne, D.T. 1993. Reproductive investment in bushcrickets: the allocation of male and female nutrients to offspring. Proc. Roy. Soc. Lond. B., 252, 1-5.
Simmons, L.W., Craig, M., Llorens, T., Schinzig, M. & Hosken, D. 1993. Bushcricket spermatophores vary in accord with sperm competition and parental investment theory. Proc. Roy. Soc. Lond. B., 251, 183-186.
Parker, G.A., Simmons, L.W. & Ward, P.I. 1993. Optimal copula duration in dungflies: effects of frequency dependence and female mating status. Behav. Ecol. Sociobiol., 32, 157-166.
Zuk, M., Simmons, L.W. & Cupp, L. 1993. Calling characteristics of parasitized and unparasitized populations of the field cricket Teleogryllus oceanicus. Behav. Ecol. Sociobiol. , 33, 339-343.
Simmons, L.W. 1994. Courtship role reversal in bushcrickets: another role for parasites? Behav. Ecol. , 5, 259-266.
Simmons, L.W. 1994. Reproductive energetics of the role reversing bushcricket, Kawanaphila nartee (Orthoptera: Tettigoniidae: Zaprochilinae). J. Evol. Biol. , 7, 189-200.
Simmons, L.W. & Zuk, M. 1994. Age structure of parasitized and unparasitized populations of the field cricket Teleogryllus oceanicus. Ethology, 333-340.
Simmons, L.W., Llorens, T., Schinzig, M., Hosken, D. & Craig, M. 1994. Sperm competition selects for male mate choice and protandry in the bushcricket, Requena verticalis (Orthoptera: Tettigoniidae). Anim. Behav., 47, 117-122.
Parker, G.A. & Simmons, L.W. 1994. Evolution of phenotypic optima: copula duration in dungflies. Nature, 370, 53-56.
Simmons, L.W. 1995. Relative parental expenditure, potential reproductive rates and the control of sexual selection in katydids. Am. Nat., 145, 797-808.
Simmons, L.W. 1995. Courtship feeding in katydids (Orthoptera: Tettigoniidae): investment in offspring and in obtaining fertilizations. Am. Nat. , 146, 307-315.
Simmons, L.W. 1995. Correlates of male quality in the field cricket, Gryllus campestris L.: age, size and symmetry determine pairing success in field populations. Behav. Ecol. , 6, 376-381.
Simmons, L.W. 1995. Male bushcrickets tailor spermatophores in relation to their remating intervals. Funct. Ecol., 9, 881-886.
Simmons, L.W., Tomkins, J.L. & Manning, J. 1995. Sampling bias and fluctuating asymmetry. Anim. Behav. 49, 1697-1699.
Tomkins, J.L. & Simmons, L.W. 1995. Patterns of fluctuating asymmetry in earwig forceps: no evidence for reliable signalling. Proc. Roy. Soc. Lond. B, 259, 89-96.
Zuk, M., Simmons, L.W. & Rotenberry, J.T. 1995. Acoustically-orienting parasitoids in calling and silent males of the field cricket Teleogryllus oceanicus. Ecol. Ent., 20, 380-383.
Simmons, L.W. & Parker, G.A. 1996. Parental investment and the control of sexual selection: can sperm competition affect the direction of sexual competition? Proc. Roy. Soc. Lond. B., 263, 515-519.
Simmons, L.W. & Ritchie, M.G. 1996. Symmetry in the songs of crickets. Proc. Roy. Soc. Lond. B., 263, 305-311.
Simmons, L.W. & Tomkins, J.L. 1996. Sexual selection and the allometry of earwig forceps. Evol. Ecol., 10, 97-106.
Simmons, L.W., Stockley, P., Jackson, R.L. & Parker, G.A. 1996. Sperm competition or sperm selection: no evidence for female influence over paternity in yellow dung flies Scatophaga stercoraria . Behav. Ecol. Sociobiol., 38, 199-206.
Allen, G.R. & Simmons, L.W. 1996. Coercive mating, fluctuating asymmetry and male mating success in the dung fly, Sepsis cynipsea. Anim. Behav. , 52, 737-741.
Parker, G.A. & Simmons, L.W. 1996. Parental investment and the control of sexual selection: predicting the direction of sexual competition. Proc. Roy. Soc. Lond. B., 263, 315-321.
Tomkins, J.L. & Simmons, L.W. 1996. Dimorphisms and fluctuating asymmetry in the forceps of male earwigs. J. Evol. Biol., 9, 753-770.
Rotenberry, J.T., Zuk, M., Simmons, L.W. & Hayes, C. 1996. Phonotactic parasitoids and cricket song structure: an evaluation of alternative hypotheses. Evol. Ecol. , 10, 233-243.
Simmons, L.W. & Kvarnemo, C. 1997. Ejaculate expenditure by male bushcrickets decreases with sperm competition intensity. Proc. Roy. Soc. Lond. B. 264, 1203-1208.
Hunt, J. & Simmons, L.W. 1997. Patterns of fluctuating asymmetry in beetle horns: an experimental examination of the honest signalling hypothesis. Behav. Ecol. Sociobiol., 41, 109-114.
Zuk, M. & Simmons, L.W. 1997. Reproductive strategies of the crickets (Orthoptera: Gryllidae). In: Choe, J.C. & Crespi, B.J. (Eds.). The Evolution of Mating Systems in Insects and Arachnids. pp 89-109. Cambridge University Press.
Simmons, L.W. & Siva-Jothy, M. 1998. Sperm competition in insects: mechanisms and the potential for selection. In: Birkhead, T. & Moller, A. (Eds.). Sexual Selection and Sperm Competition. pp. 341-434. London, Academic Press.
Hunt, J. & Simmons, L.W. 1998. Patterns of parental provisioning covary with male morphology in a horned beetle (Onthophagus taurus) (Coleoptera: Scarabaeidae). Behav. Ecol. Sociobiol., 42, 447-451.
Kvarnemo, C. & Simmons, L.W. 1998. Male potential reproductive rate influences mate choice in a bushcricket. Anim. Behav., 55, 1507-1515.
Tomkins, J.L. & Simmons, L.W. 1998. Female choice and manipulations of forceps size and symmetry in the earwig Forficula auricularia L. Anim. Behav., 56, 347-356.
Zuk, M., Rotenberry, J.T., Simmons, L.W. 1998. Calling songs of field crickets (Teleogryllus oceanicus) with and without phonotactic parasitoid infection. Evolution, 52, 166-171.
Hunt, J. & Simmons, L.W. 1998. Patterns of fluctuating asymmetry in horned beetles: no evidence for reliable signalling. Behav. Ecol. 9, 465-470.
Jones, A.G., Kvarnemo, C., Moore, G.I., Simmons, L.W., & Avise, J.C. 1998. Microsatellite evidence for monogamy and sex-biased recombination in the Western Australian seahorse Hippocampus angustus. Mol. Ecol. 7, 1497-1505.
Stockley, L.W. & Simmons, L.W. 1998. Consequences of sperm displacement for female dung flies, Scatophaga stercoraria. Proc. Roy. Soc. Lond. B. 265, 1755-1760.
Simmons, L.W., Parker, G.A. & Stockley, P. 1999. Sperm displacement in the yellow dung fly, Scatophaga stercoraria: An investigation of male and female processes. Am. Nat. 153, 302-314.
Simmons, L.W., Tomkins, J.L. & Hunt, J. 1999. Sperm competition games played by dimorphic male beetles. Proc. Roy. Soc. Lond. B 266, 145-150.
Simmons, L.W., Tomkins, J.L., Kotiaho, J.S. & Hunt, J. 1999. Fluctuating paradigm. Proc. Roy. Soc. Lond. B 266, 593-595.
Simmons, L.W., Beesley, L., Lindhjem, P., Newbound, D., Norris, J. & Wayne, A. 1999. Nuptial feeding in bushcrickets: an indicator of male quality? Behav. Ecol. 10, 263-269.
Kvarnemo, C. & Simmons, L.W. 1999. Variance in female quality, OSR and male mate choice in a bushcricket. Behav. Ecol. Sociobiol. 45, 245-252.
Rhodes, G., Roberts, J. & Simmons, L.W. 1999. Reflections on symmetry and attractiveness. Psych. Evol. Gen. 1, 279-295.
Tomkins, J.L. & Simmons, L.W. 1999. Heritability of size but not symmetry in a sexually selected trait chosen by female earwigs. Heredity 82, 151-157.
Tomkins, J.L., Simmons, L.W., Knell, R.J. & Norris, K.A. 1999. Correlates of ball size and rolling speed in the dung beetle Kheper nigroaeneus (Coleoptera: Scarabaeidae). J. Zool. 248, 483-487.
Parker, G.A., Simmons, L.W., Stockley, P. McChristie, D.M. & Charnov, E.L. 1999. Optimal copula duration in dungflies: effects of female size and egg content. Anim. Behav. 57, 795-805.
Kotiaho, J.S., Tomkins, J.L. & Simmons, L.W. 1999. Unfamiliar citations breed mistakes. Nature 400, 307.
Simmons, L.W., Tomkins, J.L. & Alcock, J. 2000. Can minor males of Dawson's burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini) compensate for reduced access to virgin females through sperm competition? Behav. Ecol. 11, 319-325.
Simmons, L.W. & Achmann, R. 2000. Microsatellite analysis of sperm utilisation patterns in the bushcricket, Requena verticalis. Evolution. 54, 942-952.
Hunt, J. & Simmons, L.W. 2000. Maternal and paternal effects on offspring phenotype in the dung beetle Onthophagus taurus. Evolution. 54, 936-941.
Parker, G.A. & Simmons, L.W. 2000. Optimal copula duration in yellow dungflies: ejaculatory duct dimensions and size-dependent sperm displacement. Evolution. 54, 924-935.
Tomkins, J.L. & Simmons, L.W. 2000. Sperm competition games played by dimorphic male beetles: fertilization gains with equal mating access. Proc. R. Soc. Lond. B. 267, 1547-1553.
Simmons, L.W. 2001. The evolution of polyandry: an examination of the genetic incompatibility and good-sperm hypotheses. J. Evol. Biol., 14, 585-594.
Simmons, L.W., Zuk, M. & Rotenberry, J.T. 2001. Geographic variation in female preference functions and male songs of the field cricket, Teleogryllus oceanicus, Evolution, 55, 1386-1394.
Hunt, J. & Simmons, L.W. 2001. Status-dependent selection in the dimorphic beetle Onthophagus taurus, Proc. R. Soc. Lond. B, 268, 2409-2414.
Kotiaho, J.S. & Simmons, L.W. 2001. Effects of Macrocheles mites on longevity of males of the dimorphic dung beetle Onthophagus binodis. J. Zool. Lond., 254, 441-446.
Kotiaho, J.S., Simmons, L.W. & Tomkins, J.L. 2001. Towards a resolution of the lek paradox. Nature, 410, 684-686.
Tomkins, J.L., Simmons, L.W. & Alcock, J. 2001. Brood provisioning strategies in Dawson's burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini). Behav. Ecol. Sociobiol., 50, 81-89.
Zuk, M., Rotenberry, J.T. & Simmons, L.W. 2001. Geographic variation in calling song of the field cricket Teleogryllus oceanicus: the importance of spatial scale. J. Evol. Biol., 14, 731-741.
Simmons, L.W. & Kotiaho, J.S. 2002. Evolution of ejaculates: Patterns of phenotypic and genotypic variation and condition dependence in sperm competition traits. Evolution, 56, 1622-1631.
Byrne, P. G, Roberts, J.D., & Simmons, L.W. 2002. Sperm competition selects for increased testis mass in Australian frogs. J. Evol. Biol., 15, 347-355.
Hunt, J. & Simmons, L.W. 2002. Behavioural dynamics of biparental care in the dung beetle Onthophagus taurus. Anim. Behav., 64, 65-75.
Hunt, J. & Simmons, L.W. 2002. The genetics of maternal care: direct and indirect genetic effects on phenotype in the dung beetle Onthophagus taurus. Proc. Natnl. Acad. Sci. USA, 99, 6828-6832.
Hunt, J. & Simmons, L.W. 2002. Confidence of paternity and paternal care: covariation revealed through the experimental manipulation of a matimg system in the beetle Onthophagus taurus. J. Evol. Biol., 15, 784-795.
Hunt, J., Simmons, L.W. & Kotiaho, J.S. 2002. A cost of maternal care in the dung beetle Onthophagus taurus? J. Evol. Biol., 15, 57-64.
Koehler, N., Rhodes, G. & Simmons, L.W. 2002. Are human female preferences for symmetrical male faces enhanced when conception is likely? Anim. Behav., 64, 233-238.
Moczek, A.P., Hunt, J., Emlen, D.J. & Simmons, L.W. 2002. Threshold evolution in exotic populations of a polyphenic beetle. Evol. Ecol. Res., 4, 587-601.
Tomkins, J.L. & Simmons, L.W. 2002. Measuring relative investment: a case study of testes investment in species with alternative male reproductive tactics. Anim. Behav., 63, 1009-1016.
Simmons, L.W. 2003. The evolution of polyandry: patterns of genotypic variation in female mating frequency, male fertilization success, and a test of the sexy-sperm hypothesis. J. Evol. Biol., 16, 624-634.
Simmons, L. W., Wernham, J., García-González, F. & Kamien, D. 2003. Variation in paternity in the field cricket Teleogryllus oceanicus: no detectable influence of sperm numbers or sperm length. Behav. Ecol., 14, 539-545.
Simmons, L.W., Alcock, J., & Reeder, A. 2003. The role of cuticular hydrocarbons in male attraction and repulsion by Dawson's burrowing bee, Amegilla dawsoni. Anim. Behav., 66, 677-685.
Byrne, P.G., Simmons, L.W. & Roberts, J.D. 2003. Sperm competition and the evolution of gamete morphology in frogs. Proc. R. Soc. Lond. B, 270, 2079-2086.
Firman, R.C., Simmons, L.W., Cummins, J.M. & Matson, P.L. 2003. Are body fluctuating asymmetry and the ratio of 2nd to 4th digit length reliable predictors of ejaculate quality? Hum. Reprod., 18, 808-812.
House, C.M. & Simmons, L.W. 2003. Genital morphology and fertilisation success in the dung beetle Onthophagus taurus: an example of sexually selected male genitalia. Proc. R. Soc. Lond. B 270, 447-455.
Kotiaho, J.S. & Simmons, L.W. 2003. Longevity cost of reproduction for males but no longevity cost of mating or courtship for females in the male-dimorphic dung beetle Onthophagus binodis. J. Insect Physiol., 49, 817-822.
Kotiaho, J.S., Simmons, L.W., Hunt, J. & Tomkins, J.L. 2003. Males influence maternal effects that promote sexual selection: a quantitative genetic experiment with dung beetles Onthophagus taurus. Am. Nat., 161, 852-859.
Rhodes, G., Chan, J., Zebrowitz, L. & Simmons, L.W. 2003. Does sexual dimorphism in human faces signal health? Proc. R. Soc. Lond. B (Suppl.), 270, S92-S95.
Tomkins, J.L. & Simmons, L.W. 2003. Fluctuating asymmetry and sexual selection: Paradigm shifts, publication bias and observer expectation. In: Developmental Stability: Causes and Consequences (Ed. by M. Polak), pp. 231-261 New York: Oxford University Press.
Simmons, L.W. 2004. Genotypic variation in calling songs and female preferences of the field cricket Teleogryllus oceanicus: common-garden and full sib analyses. Anim. Behav., 68, 313-322.
Simmons, L.W., Beveridge, M. & Krauss, S. 2004. Genetic analysis of parentage within experimental populations of a male dimorphic beetle, Onthophagus taurus, using amplified fragment length polymorphism. Behav. Ecol. Sociobiol., 57, 164-173.
Simmons, L.W., Firman, R.C., Rhodes, G. & Peters, M. 2004. Human sperm competition: testis size, sperm production, and rates of extra-pair copulations. Anim. Behav., 68, 297-302.
Simmons, L.W., Rhodes, G., Koehler, N. & Peters, M. 2004. Are human preferences for facial symmetry focused on signals of developmental stability? Behav. Ecol., 15, 864-871.
Beveridge, M. & Simmons, L.W. 2004. Microsatellite loci for Dawson's burrowing bee (Amegilla dawsoni) and their cross-utility in other Amegilla species. Mol. Ecol. Notes, 4, 379-381.
Hunt, J. & Simmons, L.W. 2004. Optimal maternal investment: application of the marginal value theorem to maternal provisioning strategies in the dung beetle Onthophagus taurus. Behav. Ecol. Sociobiol., 55, 302-312.
Koehler, N., Simmons, L.W., Rhodes, G. & Peters, M. 2004. The relationship between sexual dimorphism in human faces and fluctuating asymmetry. Proc. R. Soc. Lond. B. (Suppl.), 271, S233-S236.
Koehler, N., Simmons, L.W., Rhodes, G. 2004. How well does second to fourth digit ratio in hands correlate with other indications of masculinity in males? Proc. R. Soc. Lond. B (Suppl.), 271, S296-S298.
Kvarnemo, L. & Simmons, L.W. 2004. Testes investment is unaffected by spawning mode in pipefishes and seahorses. Biol. J. Linn. Soc., 83, 369-376.
Zuk, M., Simmons, L.W., Rotenberry, J.T. & Stoehr, A.M. 2004. Sex differences in immunity in two species of field crickets. Can. J. Zool., 82, 627-634.
Simmons, L.W. 2005. The evolution of polyandry: sperm competition, sperm selection, and offspring viability. Ann. Rev. Ecol. Evol. Syst., 36, 125-146.
Simmons, L.W. & Roberts, B. 2005. Bacterial immunity traded for sperm viability in male crickets. Science 309, 2031.
Simmons, L.W., Zuk, M. & Rotenberry, J.T. 2005. Immune function reflected in calling song characteristics in a natural population of the cricket Teleogryllus commodus. Anim. Behav. 69, 1235-1241
Alcock, J. Simmons, L.W. & Beveridge, M. 2005. Seasonal changes in offspring sex and size in Dawson's burrowing bees (Amegilla dawsoni) (Hymenoptera: Anthophorini). Ecol. Ent. 30, 247-254.
Beveridge, M. & Simmons, L.W. 2005. Microsatellite loci for the Australian field cricket Teleogryllus oceanicus and their cross-utility in Teleogryllus commodus. Mol. Ecol. Notes, 5, 733-735.
Emlen, D.J., Hunt, J. & Simmons, L.W. 2005. Evolution of sexual dimorphism and male dimorphism in the expression of beetle horns: phylogenetic evidence for modularity, evolutionary lability, and constraint. Am. Nat. 166, S42-S68.
García-González, F. & Simmons, L.W. 2005. The evolution of polyandry: intrinsic sire effects contribute to embryo viability. J. Evol. Biol. 18, 1097-1103.
García-González, F. & Simmons, L.W. 2005. Sperm viability matters in insect sperm competition. Cur. Biol. 15, 271-275.
House, C.M. & Simmons, L.W. 2005. The evolution of male genitalia: patterns of genetic variation and covariation in the genital sclerites of the dung beetle Onthophagus taurus. J. Evol. Biol., 18, 1281-1292.
House, C.M. & Simmons, L.W. 2005. Relative influence of male and female genital morphology on paternity in the dung beetle Onthophagus taurus. Behav. Ecol. 16, 889-897.
Iemmolo, M., Simmons, L.W. & Matson, P. 2005. The rapid detection of cytotoxicity using a modified human sperm survival assay. J. Assist. Reprod. Gen. 22, 177-180.
Kilgallon, S.K. & Simmons, L.W. 2005. Image content influences men's semen quality. Biol. Let. 1, 253-255.
Rhodes, G., Simmons, L.W. & Peters, M. 2005. Attractiveness and sexual behaviour: do our preferences generate variation in mating success? Evol. Hum. Behav. 26, 186-201.
Simmons, L.W. & Kvarnemo, C. 2006. Costs of breeding and their effects on the direction of sexual selection. Proc. R. Soc. Lond. B 273, 465-470.
Alcock, J., Simmons, L.W. & Beveridge, M. 2006. Does variation in female body size affect nesting success in Dawson’s burrowing bee, Amegilla dawsoni (Apidae: Anthophorini)? Ecol. Entomol. 31, 352-357.
Andersson, M. & Simmons, L.W. 2006. Sexual selection and mate choice. Trend. Ecol. Evol. 21, 296-302. (invited review for 20th anniversary issue).
Beveridge, M. & Simmons, L.W. 2006. Panmixia: an example from Dawson’s burrowing bee (Amegilla dawsoni) (Hymenoptera: Anthophorini). Mol. Ecol. 15, 951-957.
Beveridge, M. & Simmons, L.W. & Alcock, J. 2006. Genetic breeding system and investment patterns within nests of Dawson's burrowing bee (Amegilla dawsoni) (Hymenoptera: Anthophorini). Mol. Ecol. 15, 3459-3467.
House, C.M. & Simmons, L.W. 2006. Offensive and defensive sperm competition roles in the dung beetle Onthophagus taurus (Coleoptera: Scarabaeidae). Behav. Ecol. Sociobiol. 60, 131-136.
Tregenza, T., Simmons, L.W., Wedell, N. & Zuk, M. 2006. Female preference for male courtship and its role as a signal of immune function and condition. Anim. Behav. 72, 809-818.
Simmons, L.W., Beveridge, M., Wedell, N. & Tregenza, T. 2006. Post-copulatory inbreeding avoidance by female crickets only revealed by molecular markers. Mol. Ecol. 15, 3817-3824.
Simmons, L.W. & Emlen, D.J. 2006. Evolutionary trade-off between weapons and testes. Proc. Natnl. Acad. Sci. USA 103, 16346-16351.
Koehler, N., Rhodes, G., Simmons, L.W. & Zebrowitz, L.A. 2006. Do cyclic changes in women’s face preferences target cues to long-term health? Soc. Cog. 5, 641-656. Simmons, L.W. & García-González, F. 2007. Female crickets trade offspring viability for fecundity. J. Evol. Biol., 20, 1617-1623.
Simmons, L.W. & Kotiaho, J.S. 2007. The effects of reproduction on courtship, fertility and longevity within and between alternative male mating tactics of the horned beetle, Onthophagus binodis. J. Evol. Biol., 20, 488-495.
Simmons, L.W. & Kotiaho, J.S. 2007. Quantitative genetic correlation between trait and preference supports a sexually selected sperm process. Proc. Natnl. Acad. Sci. USA, 104, 16604-16608.
Simmons, L.W., Beveridge, M. & Kennington, W.J. 2007. Polyandry in the wild: temporal changes in female mating frequency and sperm competition intensity in natural populations of the tettigoniid Requena verticalis. Mol. Ecol.16, 4613-4623.
Simmons, L.W., Emlen, D.J. & Tomkins, J.L. 2007. Sperm competition games between sneaks and guards: a comparative analysis using dimorphic male beetles. Evolution, 61, 2684-2692.
Simmons, L.W., Denholm, A., Jackson, C., Levy, E. & Madon, E. 2007. Male crickets adjust ejaculate quality with both risk and intensity of sperm competition. Biol. Lett. 3, 520-522.
García-González, F. & Simmons, L.W. 2007. Paternal indirect genetic effects on offspring viability and the benefits of polyandry. Curr. Biol. 17, 32-36.
García-González, F. & Simmons, L.W. 2007. Shorter sperm confer higher competitive fertilization success. Evolution, 61, 816-824.
House, C.M. & Simmons, L.W. 2007. No evidence for condition-dependent expression of male genitalia in the dung beetle Onthophagus taurus. J. Evol. Biol., 20, 1322-1332.
Peters, M., Rhodes, G. & Simmons, L.W. 2007. Contributions of the face and body to overall attractiveness. Anim. Behav. 73, 937-942.
Rhodes, G. & Simmons, L.W. 2007. Symmetry, attractiveness, and sexual selection. In: Oxford Handbook of Evolutionary Psychology (Eds. Barrett, L. & Dunbar, R.). Oxford University Press, Oxford. pp. 333-364.
Rhodes, G., Yoshikawa, S., Palermo R., Simmons, L.W., Peters, M., Lee, K., Halberstadt, J. & Crawford, J.R. 2007. Perceived health contributes to the attractiveness of facial symmetry, averageness and sexual dimorphism. Perception, 36, 1244-1252.
Thomas, M.L. & Simmons, L.W. 2007. Male crickets adjust the viability of their sperm in response to female mating status. Am. Nat. 170, 190-195.
Simmons, L.W. & García-González, F. 2008. Evolutionary reduction in testes size and competitive fertilization success in response to the experimental removal of sexual selection in dung beetles. Evolution, 62, 2580-2591.
Simmons, L.W. & Thomas, L.W. 2008. No postcopulatory response to inbreeding by male crickets. Biol. Lett., 4, 183-185.
Simmons, L.W., Beveridge, M. & Evans, J.P. 2008. Molecular evidence for multiple paternity in a feral population of green swordtails. J. Hered. 99, 610-615.
Simmons, L.W. & Emlen, D.J. 2008. No fecundity cost of female secondary sexual trait expression in the horned beetle Onthophagus sagittarius. J. Evol. Biol., 21, 1227-1235.
Cotter, S.C., Beveridge, M. & Simmons, L.W. 2008. Male morph predicts investment in larval immune function in the dung beetle, Onthophagus taurus. Behav. Ecol., 19, 331-337.
Dziminski, M. A., Roberts, J.D. & Simmons, L.W. 2008. Fitness consequences of parental compatibility in the frog Crinia georgiana. Evolution, 62, 879-886.
Evans, J.P. & Simmons, L.W. 2008. The genetic basis of traits regulating sperm competition and polyandry: can selection favour the evolution of good- and sexy-sperm? Genetica, 134, 5-19.
Firman, R.C. & Simmons, L.W. 2008. Polyandry facilitates post-copulatory inbreeding avoidance in house mice. Evolution, 62, 603-611.
Firman, R.C. & Simmons, L.W. 2008. Polyandry, sperm competition and reproductive success in mice. Behav. Ecol., 19, 695-702.
Firman, R.C. & Simmons, L.W. 2008. The frequency of multiple paternity predicts variation in testes size among island populations of house mice. J. Evol. Biol., 21, 1524-1533.
Humphries, S., Evans, J.P. & Simmons, L.W. 2008. Sperm competition: linking form to function. BMC Evolutionary Biology, 8, 319. doi:10.1186/1471-2148-8-319.
Lie, H.C., Rhodes, G., & Simmons, L.W. 2008. Genetic diversity revealed in human faces. Evolution, 62, 2474-2486.
Peters, M., Rhodes, G. & Simmons, L.W. 2008. Does attractiveness in men provide clues to semen quality? J. Evol. Biol., 21, 572-579.
Peters, M., Simmons, L.W. & Rhodes, L.W. 2008. Testosterone is associated with mating success but not attractiveness or masculinity in human males. Anim. Behav., 76, 297-303.
Thomas, M. & Simmons, L.W. 2008. Cuticular hydrocarbons are heritable in the cricket Teleogryllus oceanicus. J. Evol. Biol., 21, 801-806.
Thomas, M. & Simmons, L.W. 2008. Rival male relatedness does not affect ejaculate allocation as predicted by sperm competition theory. PLoS One, 3, e2151. doi:10.1371/journal.pone.0002151
Thomas, M. & Simmons, L.W. 2008. Sexual dimorphism in cuticular hydrocarbons of the Australian field cricket Teleogryllus oceanicus (Orthoptera: Gryllidae). J. Insect Physiol., 1081-1089.
Wedell, N., Tregenza, T. & Simmons, L.W. 2008. Nuptial gifts fail to resolve a sexual conflict in an insect. BMC Evol. Biol., 8, 204. doi: 10.1186/1471-2148-8-204
Werner, M. & Simmons, L.W. 2008. The evolution of male genitalia: functional integration of genital sclerites in the dung beetle Onthophagus taurus. Biol. J. Linn. Soc., 93, 257-266. Werner, M. & Simmons, L.W. 2008. Insect sperm motility. Biol. Rev., 83, 191-208.
Simmons, L.W. & Moore, A.J. 2009. Evolutionary quantitative genetics of sperm. In: Birkhead, T.R., Hosken, D.J. & Pitnick, S. (eds.) Sperm Evolution. pp. 405-434. Elsevier, London.
Simmons, L.W., Roberts, J.D. & Dziminski, M.A. 2009. Egg-jelly influences sperm motility in the externally fertilizing frog, Crinia georgiana. J. Evol. Biol., 22, 225-229.
Simmons, L.W., House, C.M., Hunt, J. & García-González, F. 2009. Evolutionary response to sexual selection in male genital morphology. Curr. Biol. 19, 1442-1446. DOI 10.1016/j.cub.2009.06.056
Alcock, J., Bailey, W.J., & Simmons, L.W. 2009. The Mating System of Amegilla (Asarapoda) paracalva Brooks (Hymenoptera: Apidae). J. Insect Behav.
Dowling, D.K. & Simmons, L.W. 2009. ROS as universal constraints in life-history evolution. Proc. R. Soc. Lond. B., 276, 1737-1745.
Dziminski, M.A., Roberts, J.D. & Simmons, L.W. 2009. Sperm competitiveness in frogs: slow and steady wins the race. Proc. R. Soc. Lond. B., 276, 3955-3961.
Dziminski, M.A., Roberts, J.D. & Simmons, L.W. 2009. Sperm morphology, motility, and fertilization capacity in the myobatrachid frog Crinia georgiana. Reprod. Fert. Dev.
Firman, R.C. & Simmons, L.W. 2009. Sperm competition and the evolution of the sperm hook in house mice. J. Evol. Biol.
Firman, R.C. & Simmons, L.W. 2009. Experimental evolution of sperm quality via postcopulatory sexual selection in house mice. Evolution.
Lie, H.C., Simmons, L.W. & Rhodes, G. 2009. Does genetic diversity predict health in humans? PLoS One, 4(7), e6391. doi:10.1371/journal.pone.0006391
Lie, H.C., Simmons, L.W. & Rhodes, G. 2009. Genetic dissimilarity, genetic diversity, and mate preferences in humans. Evol. Hum. Behav.
Peters, M., Simmons, L.W., & Rhodes, G. 2009. Photographs of male faces and bodies provide no evidence for a preference shift for masculinity or symmetry across the menstrual cycle. PLoS One, 4, e4138. doi:10.1371/journal.pone.0004138.
Polak, M. & Simmons, L.W. 2009. Secondary sexual trait size reveals competitive fertilization success in Drosophila bipectinata Duda. Behav. Ecol., 20, 753-760.
Robinson, S.P., Kennington, W.J. & Simmons, L.W. 2009. No evidence for optimal fitness at intermediate levels of inbreeding in Drosophila melanogaster. Biol. J. Linn. Soc., 98, 501-510.
Thomas, M.L. & Simmons, L.W. 2009. Male-derived cuticular hydrocarbons signal sperm competition intensity and affect ejaculate expenditure in crickets. Proc. R. Soc. Lond. B., 276, 383-388.
Thomas, M.L. & Simmons, L.W. 2009. Sexual selection on cuticular hydrocarbons in the Australian field cricket, Teleogryllus oceanicus. BMC Evol. Biol. 9, 162. doi:10.1186/1471-2148-9-162
Thomas, M.L. & Simmons, L.W. 2009. Male dominance influences pheromone expression, ejaculate quality, and fertilization success in the Australian field cricket, Teleogryllus oceanicus. Behav. Ecol. 20, 1118-1124.
Wojcieszek, J.M. & Simmons, L.W. 2009. Isolation and characterisation of 11 polymorphic microsatellite loci in the millipede Antichiropus variabilis Attems (Diplopoda: Polydesmida: Paradoxosomatidae). Mol. Ecol. Res., 9, 1208-1211.
Young, M.J., Simmons, L.W. & Evans, J.P. 2009. Isolation and characterization of 12 novel DNA microsatellites in the Western Rainbowfish, Melanotaenia australis. Mol. Ecol. Res., 9, 1252-1254.
Books
Simmons, L.W. 2001. Sperm Competition and its Evolutionary Consequences in the Insects. Princeton University Press, Princeton.
Lucus, J.R. & Simmons, L.W. (Eds.) 2006. Essays in Animal Behaviour. Celebrating 50 years of Animal Behaviour. Elsevier, London.
Other publications
Simmons, L.W. 2001. A Darwinian perspective on fertility. Assisted Reproductive Technology and Science, 1, 4-5.
Lessells, C. (K) L., Bennet, A.T.D., Birkhead, T.R., Colegrave, N., Dall, S.R.X., Harvey, P.H., Hatchwell, B., Hosken, D.J., Hunt, J., Moore, A.J., Parker, G.A., Pitnick, S., Pizzari, T., Radwan, J., Ritchie, M., Sheldon, B.C., Simmons, L.W., Stockley, P., Tregenza, T. & Zuk, M. 2006. Debating sexual selection and mating systems. Science, 689-690.
Pizzari, T., Birkhead, T.R., Blows, M.W., Brooks, R., Buchanan, K.L., Clutton-Brock, T.H., Harvey, P.H., Hosken, D.J., Jennions, M.D., Kokko, H., Kotiaho, J.S., Lessells, C. (K) L., Marcias-García, C., Moore, A.J., Parker, G.A., Partridge, L., Pitnick, S., Radwan, J., Ritchie, M., Sheldon, B.C., Simmons, L.W., Snook, R.R., Stockley, P. & Zuk, M. 2006. Debating sexual selection and mating systems. Science, 690. Ridsdill-Smith, J. & Simmons, L.W. 2009. Dung Beetles. In Resh, V. H. & Cardé, R.T. Encyclopedia of Insects. pp 304-307. Elsevier Science, USA.
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